Captain America in last line
Scotland defeated Japan on Wednesday to move to the top of Pool B ahead of the encounter at Elland Road in Leeds.
Try-scorer in the 16-25 loss to Samoa, Wyles resumes his role at fullback following two consecutive matches on the wing, moving Blaine Scully to the touchline opposite Taku Ngwenya.
Phil Thiel gets a start at hooker, replacing Zach Fenoglio, who helped provide a stable scrum for 50 minutes. Chris Baumann scored his first international try against Samoa in front of a pro-U.S. crowd at Brighton Community Stadium, and will be available as front row cover alongside Fenoglio and Olive Kilifi.
"The scrum was once again a good platform of attack and the line-out had a few hiccups, which was unfortunate as it cost us on four occasions [against Samoa]," Tolkin said. "Fenoglio's logged a lot of minutes as starter [this year] and Thiel is a little fresher.
"They are both very good and virtually even, so we thought this was a good time to get Phil in as a starter and have Zach add punch in off the bench."
The big bodies of Danny Barrett, Cam Dolan, and John Quill also provide Tolkin with options from the bench, while the creativeness of Folau Niua and Shalom Suniula could open a tired Scotland defense.
The two-try output against Samoa bests the previous matchup in San Jose earlier this summer, but the Eagles were unable to dot down for a five-pointer last year against their upcoming opponent Scotland.
Wyles kicked two penalties as the humid Texas heat kept a fluid attack from forming throughout the match. For the victors, scrumhalf and captain Greig Laidlaw converted all three Scotland tries and successfully kicked a penalty.
"We have the capability to do some dynamic things with ball in hand, with both power and pace," Tolkin said. "We'd like to be able to make this a more consistent, potent weapon. Scotland, like everyone, has weaknesses and we need to exploit them effectively with both of those tools.
"Last summer, we played them first up in the test series with many players who had not played much rugby prior to that match, so we didn't have attacking rhythm and confidence, which we showed more of in the next two matches," he added.
USA: 15 Chris Wyles (captain), 14 Takudzwa Ngwenya, 13 Seamus Kelly, 12 Thretton Palamo, 11 Blaine Scully, 10 AJ MacGinty, 9 Mike Petri, 8 Samu Manoa, 7 Andrew Durutalo, 6 Al McFarland, 5 Greg Peterson, 4 Hayden Smith, 3 Titi Lamositele, 2 Phil Thiel, 1 Eric Fry.
Replacements: 16 Zach Fenoglio, 17 Oli Kilifi, 18 Chris Baumann, 19 Cameron Dolan, 20 John Quill, 21 Danny Barrett, 22 Shalom Suniula, 23 Folau Niua.
Date: Sunday, September 27
Venue: Elland Road, Leeds
Kick-off: 14.30 (13.30 GMT, 09.30 EDT)
Referee: Chris Pollock (New Zealand)
Assistant referees: John Lacey (Ireland), Mike Fraser (New Zealand)
TMO: Ben Skeen (New Zealand)
